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Alternate story segments may focus on Chris Redfield and his new partner Jessica Sherawat, or new BSAA members Keith and Quint.

Jill Valentine works with her new partner Parker Luciani for the BSAA (Bioterrorism Security Assessment Alliance). The story is broken into chapters fitting for a portable title story-wise, chapters alternate timelines. The primary focus of the story is on Jill Valentine and Chris Redfield as they hunt for answers on a mysterious luxury cruise ship known as the Queen Zenobia. The game takes place before part five in 2005, and after the total destruction of the ultra-modern sea-based city of Terragrigia by terrorists using old Umbrella-style B.O.W.s, or Bio-Organic Weapons. Resident Evil: Revelations marks a return to the survival horror roots of the franchise from the action-centric gameplay of the previous title, Resident Evil 5.